The Archaeognatha are an order of wingless insects, also known as jumping bristletails. They are among the least evolutionarily changed insects, appearing in the Middle Devonian period along with the arachnids. They are known from both body and trace fossils (the latter including body imprints and trackways) throughout the remainder of the Paleozoic Era.[2] The name Archaeognatha is derived from Greek, archaeos meaning "ancient" and gnatha meaning "jaw". This refers to the articulation of the mandibles, which has a single condyle, where all higher insects have two. An alternative name, Microcoryphia, comes from the Greek micro, meaning "small", and coryphia, meaning head.
The order Archaeognatha has previously been combined with the order Thysanura; both orders possess three-pronged tails comprising two cerci and an epiproct.
The approximately 350 species in the two families are distributed worldwide. No species is currently at conservation risk.
Description
Archaeognaths are small insects with elongated bodies bent into an arch shape and three long tail-like structures, of which the median filament is longest. They have flexible antennae, large compound eyes that meet at the top of the head, and three ocelli. Their mouthparts are partially retractable, with simple chewing mandibles and long maxillary palps.[1]:341–343
Archaeognatha differ from Thysanura in being able to use their tails to spring up to 30 cm (12 in) into the air. They are also unique in possessing small, articulated "styli" on the hind (and sometimes middle) coxae and sternites 2 to 9, which are sometimes considered to be rudimentary appendages, as well as paired eversible membranous vesicles through which they absorb water.
Example of compound eyes (Trigoniophthalmus alternatus)
Further unusual features are that the abdominal sternites are each composed of three sclerites, and they cement themselves to the substrate before molting. Like Thysanura, the body is covered with scales, with a thin exoskeleton that is susceptible to dehydration.